Scientific Repository

Dinos101

Explore each species discovered by paleontology: where it lived, which bones were found, and how our understanding evolved over time.

101
Curated species
3
Periods
22
Countries
6
Brazilian species
Filter by: 101 species
Chasmosaurus belli

CA · 76.5–75 Ma

Chasmosaurus

Chasmosaurus belli

"Bell's opening lizard"

Chasmosaurus belli is the prototype ceratopsid of the subfamily Chasmosaurinae, which bears its name. It lived in the late Campanian of the Cretaceous, approximately 76.5 to 75 million years ago, in Alberta, Canada, and is one of the first large ceratopsids described by science. Its most distinctive feature is the parieto-squamosal frill that is extraordinarily large relative to the skull, with two large oval openings (the parietal fenestrae) that substantially reduced the weight of the structure. The genus name Chasmosaurus refers precisely to these openings: 'khasma' in Greek means opening or chasm. This long frill, which could exceed 60% of the total skull length, is the morphological signature of the entire subfamily. The animal was originally described by Lawrence Lambe in 1902 based on fragmentary material from the Dinosaur Park Formation, but the species was only definitively established by Lambe in 1914. Subsequent studies by Brown and Schlaikjer (1940) and Dodson (1990, 1996) substantially refined the anatomy and systematics of the taxon. Chasmosaurus belli is distinguished from other species in the genus (such as C. russelli and C. irvinensis, now reclassified as Vagaceratops) mainly by the dimensions and proportions of the frill and by the relatively short facial horns. The nasal horn is low and laterally compressed, while the supraorbital horns range from short to moderate. In 2010, a juvenile specimen of Chasmosaurus belli was discovered by Brian Campione at Dinosaur Provincial Park and studied by Phillip Bell and colleagues. The importance of this fossil goes beyond the rarity of juveniles in ceratopsids: it preserved mummified skin impressions in different regions of the body, including large hexagonal scales on the flanks, smaller scales on the neck and face, and scales organized in a differentiated pattern around the eye. Pigmentation analyses preserved through scanning electron microscopy revealed melanosomes suggesting contrasting pigmentation, possibly with a counter-shading pattern (dark dorsum, pale ventrum), which would be the first direct evidence of coloration in a ceratopsid. Chasmosaurus belli's position as the founding taxon of Chasmosaurinae makes it fundamental for understanding the evolution of the entire subfamily. Modern phylogenetic analyses consistently position C. belli as one of the most basal chasmosaurines, from which more derived forms with even more elaborate frills radiate, including Pentaceratops, Anchiceratops, Torosaurus, and Triceratops. The fossil record of Chasmosaurus belli in the Dinosaur Park Formation is one of the richest of any ceratopsid, with dozens of specimens collected over more than a century of systematic excavations.

Cretáceous Herbivore 4.9m
Confuciusornis sanctus

CN · 125–120 Ma

Confuciusornis

Confuciusornis sanctus

"Sacred Confucius bird"

Confuciusornis sanctus is the most abundant Mesozoic bird in the world fossil record: hundreds of specimens, many exquisitely preserved with feathers, have been extracted from the Yixian and Jiufotang formations of Liaoning, northeastern China. This unparalleled abundance made Confuciusornis the most studied extinct bird taxon in the world, capable of providing data on morphology, ontogeny, coloration, paleoecology, and social behavior simply not accessible for rarer birds. It lived during the Barremian to Aptian of the Early Cretaceous, approximately 125 to 120 million years ago, and represents one of the most important morphological innovations in bird history: it was the first toothless Mesozoic bird with a true horny beak and a fused tail bone forming a pygostyle. Confuciusornis morphology combines primitive and derived features remarkably. On one hand, the forelimbs retain three functional clawed fingers, and the skull retains some primitive skeletal features relative to modern birds. On the other, the complete absence of teeth, the horny beak, and the pygostyle are innovations we know mainly from modern birds, making Confuciusornis much more externally similar to a modern bird than Archaeopteryx or other contemporaries. The tail, short due to the pygostyle, accommodated in long-tailed forms a pair of very long, narrow ribbon-like rectrices whose function was probably social display. The presence or absence of these tail feathers in different specimens has historically been interpreted as sexual dimorphism, but recent analyses with ontogenetically comparable specimens question this simple interpretation. Confuciusornis flight is considered truly active, not just gliding: the sternum morphology, with well-developed keel for flight muscle insertion, and wing proportions suggest sustained powered flight capability. However, biomechanical analyses indicate the flight pattern would differ from modern birds: the shoulder joint, clavicle position, and relative proportions of humerus and ulna suggest an aerodynamically less efficient flight style compared to neornithean birds. Bone histology studies show Confuciusornis had slower growth rates than modern birds of comparable size, reaching skeletal maturity in one or two years, closer to reptiles than to modern birds.

Cretáceous Omnivore 0.5m
Dracorex hogwartsia

US · 66–66 Ma

Dracorex

Dracorex hogwartsia

"Dragon king of Hogwarts"

Dracorex hogwartsia is one of the dinosaurs with the most peculiar story in paleontology: it was named by children, in honor of a fictional school of witchcraft, and its status as a valid species is one of the most contentious taxonomic debates of the Late Cretaceous. Described in 2006 by Robert Bakker and collaborators based on a nearly complete skull discovered in the Hell Creek of South Dakota by a group of amateur fossil hunters, the animal presented a cranial morphology radically different from other known pachycephalosaurids: the skull was long, low, and flat, without any trace of a bony dome, but decorated with an impressive series of horns and nodules along the frontoparietal and temporal region. The visual result is, in fact, remarkably similar to medieval descriptions of dragons, which inspired the specimen's donors, children from the Children's Museum of Indianapolis, to propose the name to the paleontologist. The main scientific controversy surrounding Dracorex was raised in 2009, when Jack Horner and Mark Goodwin published a detailed histological analysis of the skull. Upon examining the microstructure of the frontoparietal bone, they found immature bone tissue (fibrolamellar, with abundant vasculature and without complete remodeling) characteristic of young individuals, not adults with completed bone growth. Horner and Goodwin's hypothesis is that Dracorex hogwartsia is not a distinct species, but rather a juvenile of Pachycephalosaurus wyomingensis: the flat skull with horns would be the initial state, and the dome would form progressively during growth, similar to the development of horns in sheep. This interpretation would also unify Stygimoloch spinifer as a teenager of Pachycephalosaurus, consolidating three names into a single taxon. The synonymy proposal, while widely accepted by many specialists, remains formally controversial. Other researchers, such as Robert Sullivan (2006) and David Evans and collaborators (2013), argue that the morphological differences between the skulls are too pronounced to be explained by ontogeny alone, and that Dracorex may represent a distinct lineage of flat-skulled pachycephalosaurids that lived in sympatry with Pachycephalosaurus. The question is complicated by the rarity of specimens: only one Dracorex skull is known with certainty, making it impossible to establish complete ontogenetic series. Regardless of taxonomic resolution, the Dracorex skull is one of the most extraordinary known among ornithischian dinosaurs. The tubercles, spines, and nodules covering the cranial surface have no parallel in any other pachycephalosaurid, and their function continues to be debated: they may have served for intraspecific recognition, surface thermoregulation, sexual display, or passive defense. The preservation of the specimen, with unusually well-conserved cranial surface details, has made Dracorex one of the most studied and artistically represented dinosaurs of the last twenty years.

Cretáceous Herbivore 2.4m
Euoplocephalus tutus

CA · 76.5–66 Ma

Euoplocephalus

Euoplocephalus tutus

"Well-armored and protected head"

Euoplocephalus tutus is the best-documented ankylosaur in the history of paleontology: more than 40 specimens have been collected since the original description in 1902, including several complete skulls, dermal armor in association, and at least one complete tail with preserved bony club. It lived in the late Campanian of the Cretaceous, approximately 76.5 to 66 million years ago, in North America, with records in Alberta (Canada) and possibly Montana (USA). It is the only ankylosaur for which sufficient material exists to characterize individual and ontogenetic morphological variation within the species with any degree of confidence. The animal had extensive dermal armor composed of osteoderms of multiple types: large flat plates on the back, cones or spines along the flanks, smaller scales filling the spaces between larger ones, and a hardened skin covering over the skull. Notably, Euoplocephalus had ossified eyelids, a unique adaptation protecting the eyes from predatory attacks. The tail terminated in a bony club (fused tail osteoderm, the 'golf club') whose size and robustness were analyzed by Arbour (2009) as sufficient to generate impact forces capable of fracturing bones of large predators like Tyrannosaurus and Gorgosaurus. In terms of systematics, the history of Euoplocephalus is complex. Many ankylosaur specimens from Alberta were originally referred to E. tutus based on the assumption that there would be only one ankylosaur species per formation. More recent research by Arbour and Currie (2013, 2015) reassessed this material and concluded that several specimens previously referred to E. tutus belong to distinct genera and species (such as Scolosaurus cutleri, Anodontosaurus lambei, and Dyoplosaurus acutosquameus). After this revision, the number of specimens truly referable to E. tutus decreased, but the taxon remains the best-characterized ankylosaur of the Campanian of North America. Arbour's (2009) biomechanical analysis of the tail club is one of the most cited results in ankylosaur paleobiology: using structural mechanics models and comparisons with modern hammers, Arbour demonstrated that the E. tutus club could generate impact forces of 2-6 kN, sufficient to fracture the ribs or tibiae of Tyrannosaurus or Gorgosaurus. Tail musculature, inferred from well-developed transverse processes, was capable of moving the club in high-speed lateral arcs. This analysis changed the scientific understanding of the tail club from a possible display ornament to an active and effective defensive weapon.

Cretáceous Herbivore 6m
Giraffatitan brancai

TZ · 154–150 Ma

Giraffatitan

Giraffatitan brancai

"Giraffe titan"

Giraffatitan brancai was a giant brachiosaurid sauropod that lived during the Kimmeridgian to Tithonian of the Late Jurassic, approximately 154 to 150 million years ago, in what is now Tanzania. At 26 meters in length and estimated body mass of 35,000 to 40,000 kg, it was one of the largest land animals of its time. Giraffatitan's morphology is characterized by forelimbs significantly longer than the hindlimbs, resulting in a forward-sloping back and an extremely long and high neck that allowed the animal to reach the vegetation of tree canopies in the East African Jurassic forest. Unlike many sauropods that held the neck horizontally, Giraffatitan's skeletal proportions indicate the neck was raised at a steep angle, giving the animal a vertical silhouette reminiscent of a giraffe, hence the name. The separation of Giraffatitan from Brachiosaurus as a distinct genus was proposed by Paul (1988) and confirmed by Taylor (2009) after detailed cladistic analysis. Key anatomical differences include skull configuration, taller and with a more prominent nasofrontal crest in Giraffatitan, and cervical vertebrae proportions, more elongate and with lower robustness index in Giraffatitan than in Brachiosaurus altithorax from North America. The validity of Giraffatitan as a separate genus is currently consensus in the paleontological community, though some authors have suggested both could belong to the same genus with subgenus status. Fossil material of Giraffatitan brancai is exceptionally rich for a giant sauropod: German expeditions to Tanzania between 1909 and 1913, led by Werner Janensch, recovered remains of dozens of individuals, including materials from at least five partially articulated individuals. The composite specimen HMN SII, mounted at the Museum für Naturkunde in Berlin, is the tallest mounted dinosaur skeleton in the world, at 13.27 meters. The logistical operation of the Tendaguru expeditions was one of the largest paleontological endeavors in history: more than 250 African workers transported hundreds of tons of fossilized material over 60 kilometers from the East African coast under extremely adverse conditions.

Jurassic Herbivore 26m
Heterodontosaurus tucki

ZA · 201–196 Ma

Heterodontosaurus

Heterodontosaurus tucki

"Tuck's different-toothed lizard"

Heterodontosaurus tucki is one of the most primitive ornithischians known and certainly the most anatomically surprising of its time. It lived in the Early Jurassic, approximately 201 to 196 million years ago, on the semi-arid plains of what is now South Africa, in the Upper Elliot Formation. At only 1.2 meters in length and under 2 kg, it was an agile biped with long hind limbs and relatively robust forelimbs, likely using its hands to manipulate vegetation or small prey. Its skeleton is extraordinarily well preserved, with the skull SAM-PK-K1332 being one of the most complete among primitive Triassic-Jurassic dinosaurs. The defining feature of the genus, reflected in its name, is its radically heterogeneous dentition: unlike the vast majority of herbivorous dinosaurs, Heterodontosaurus possessed three distinct types of teeth. At the front of the upper jaw were small incisor-like teeth for cropping vegetation, followed by large canine-like tusks at the corners of the maxilla and dentary, and taller, laterally compressed cheek teeth with crushing edges for processing plant food. This combination of three distinct dental morphologies in a single animal is unique among Dinosauria and generated decades of debate about function and diet. The question of the tusks is the most controversial aspect of Heterodontosaurus biology. Crompton and Charig, when describing the species in 1962, suggested omnivorous use. Later studies, especially those of Santa Luca (1980) and Sereno (2012), raised the hypothesis of sexual dimorphism: the tusks would be exclusive to males and would function in display or intraspecific combat, analogous to modern deer and suids. Norman et al. (2011), upon reexamining the material, concluded that the canine teeth were not related to carnivory but rather to social behavior. Porro et al. (2011) performed biomechanical analysis of the jaw and confirmed that its structure was highly adapted to processing hard plant material. Heterodontosaurus is also notable for its phylogeny. For decades it was placed at the base of ornithopods, but modern analyses position it at the base of Ornithischia or as a member of a lineage sister to Genasauria. This makes it a valuable window into the early stages of ornithischian dinosaur evolution, before the great diversification that produced hadrosaurs, ceratopsians, ankylosaurs, and stegosaurs. Additionally, possible evidence of keratin on the anterior teeth suggests a partial horny beak, as in many modern birds and turtles.

Jurassic Omnivore 1.2m
Kosmoceratops richardsoni

US · 76.4–75.5 Ma

Kosmoceratops

Kosmoceratops richardsoni

"Richardson's ornate horned face"

Kosmoceratops richardsoni is the ceratopsian with the most elaborate cranial ornamentation ever documented in dinosaur history: a total of 15 horns and bony structures on the skull, including a large downward-curved nasal horn, two supraorbital horns above the eyes, two jugal horns, two laterally-directed epijugal horns, and ten forward-curving apical processes on the parieto-squamosal frill. No other known dinosaur or fossil vertebrate has such dense and diversified cranial ornamentation. The animal lived approximately 76 to 75.5 million years ago during the late Campanian stage of the Cretaceous, in the region corresponding to the present-day state of Utah in the United States. Kosmoceratops belonged to the fauna of the southern portion of Laramidia Island, a large continental island that existed during the Late Cretaceous when the Western Interior Seaway divided North America into two landmasses. Notably, the southern and northern portions of Laramidia were separated by a shallow internal sea and presented completely distinct dinosaur faunas: while the south (where Kosmoceratops lived) had short-frilled ceratopsians with elaborate ornamentation, the north produced forms with longer frills. This geographic faunal difference was one of the central arguments presented by Sampson et al. (2010) in the paper describing the species. The holotype and referred specimens were collected from the Kaiparowits Formation in southern Utah, one of the most productive Late Cretaceous geological units in North America. The formation dates from the late Campanian (~76.6 to 74.5 Ma) and preserves a rich subtropical ecosystem with multiple species of dinosaurs, crocodilians, turtles, lizards, mammals, and amphibians. Kosmoceratops fossils were collected by the Utah Museum of Natural History between 2004 and 2006 at sites within the Grand Staircase-Escalante National Monument. The function of the extravagant cranial ornamentation is a subject of ongoing scientific debate. Sampson et al. (2010) argue that the structures were primarily for intraspecific display and species recognition, analogous to the horns of modern antelopes and cervids. The fact that the supraorbital horns and frill processes face laterally and forward rather than forward in combat position supports the display hypothesis over the defensive one. Later biomechanical analyses by Mallon and Anderson (2013) suggest that ceratopsids used their horns in ritualized intraspecific combat, not as defense against large predators.

Cretáceous Herbivore 4.5m
Mussaurus patagonicus

AR · 221–205 Ma

Mussaurus

Mussaurus patagonicus

"Mouse lizard from Patagonia"

Mussaurus patagonicus is a Late Triassic sauropodomorph that stars in one of paleontology's most unusual stories: it was named the 'mouse lizard' because the first discovered specimens were tiny hatchlings only 20 centimeters long, small enough to fit in the palm of a hand. When José Bonaparte and Martín Vince described the species in 1979, based on these hatchlings from the Los Colorados Formation of Argentine Patagonia, they imagined a modestly sized adult. Decades later, when adult specimens were found and described in detail, it was revealed that Mussaurus grew to 6 meters in length and 1.5 tonnes: one of the largest terrestrial animals of the Late Triassic in South America. This discrepancy between hatchlings and adults is biologically significant. It demonstrates that basal sauropodomorphs underwent dramatic ontogenetic changes not only in size but also in locomotor posture. Mussaurus hatchlings were facultative bipeds, with proportionally similar fore and hind limbs, while adults had more robust forelimbs and likely transitioned to a quadrupedal posture upon reaching large sizes. This pattern echoes what later became obligatory in the giant sauropods of the Jurassic and Cretaceous. The most impactful publication on Mussaurus in recent years was the 2021 study by Otero et al. published in Scientific Reports. Analysis of multiple nests with eggs, neonatal hatchlings, and spatially grouped juveniles revealed strong evidence of gregarious behavior: the animals gathered in age-segregated groups, suggesting some form of parental care or structured social behavior. Mussaurus eggs were preserved in communal nests at a depth suggesting they were buried for incubation, similar to the behavior of crocodiles and some extant birds. This finding pushes the origin of gregarious behavior in sauropodomorphs back to the Triassic, more than 50 million years earlier than previously thought. Phylogenetically, Mussaurus occupies a key position in the transition between solitary basal sauropodomorphs and the large gregarious sauropods of the Mesozoic. The most recent analyses place it within Massopoda, near the base of the clade that will give rise to true Sauropoda. Its body still preserves plesiomorphic features such as a moderately long neck, more robust hind limbs than forelimbs, and simple leaf-shaped teeth, but already shows derived features such as elongated cervical vertebrae and a sauropod-like tarsal formula. The exceptional fossil record of the Los Colorados Formation, with multiple age classes preserved together, makes Mussaurus the most important Triassic sauropodomorph for studies of growth, ontogeny, and social behavior.

Triassic Herbivore 6m
Tenontosaurus tilletti

US · 115–108 Ma

Tenontosaurus

Tenontosaurus tilletti

"Tillett's sinew lizard"

Tenontosaurus tilletti was a medium-sized iguanodontian ornithopod that lived during the Aptian-Albian of the Early Cretaceous, approximately 115 to 108 million years ago, in what is now the western United States. At 6 to 7.5 meters in length and estimated body mass of 700 to 1,000 kg, it was one of the most common and abundant herbivores of its time and region. The genus is morphologically distinct from other ornithopods by its extraordinarily long and robust tail, which in adults could represent more than half the total length of the animal. The caudal vertebrae are reinforced by ossified tendons, hence the name 'sinew lizard', which provided rigidity to the tail and possibly functioned as a counterweight to maintain balance during bipedal or quadrupedal locomotion. Tenontosaurus was capable of moving both on two and four legs depending on speed and terrain. The paleontological importance of Tenontosaurus extends far beyond its intrinsic morphology: the animal is the center of the longest and most influential debate in paleontology about cooperative predatory behavior in dinosaurs. When John Ostrom described Deinonychus antirrhopus in 1969 and 1970, he based much of his argument about pack hunting on the recurring spatial association of multiple Deinonychus teeth with Tenontosaurus carcasses in the Cloverly Formation. Ostrom inferred that groups of Deinonychus cooperatively attacked much larger Tenontosaurus individuals, similarly to the behavior of modern lions. This hypothesis became enormously influential and directly inspired the representation of Velociraptors as cooperative hunters in Jurassic Park (1993). However, subsequent analyses questioned the cooperative hunting interpretation. Roach and Brinkman (2007) argued that the association of Deinonychus with Tenontosaurus carcasses is more consistent with competitive scavenging behavior, like that observed in modern Komodo dragons feeding on the same carcass without true cooperation and frequently attacking each other during the feast. Evidence of multiple dead Deinonychus at the same localities as Tenontosaurus would suggest Tenontosaurus was capable of actively defending itself, killing some of its attackers. This debate about cooperative hunting versus competitive feeding in Deinonychus, centered on Tenontosaurus carcasses, remains without definitive resolution and continues to be one of the most stimulating problems in dinosaur behavior.

Cretáceous Herbivore 6.5m
Yi qi

CN · 163–159 Ma

Yi qi

Yi qi

"Strange wings"

Yi qi (pronounced approximately 'ee chee') was a diminutive scansoriopterygid dinosaur that lived during the Middle to Late Jurassic, Oxfordian stage, approximately 163 to 159 million years ago, in the region of present-day Hebei, northern China. At only 0.6 meters in length with an estimated body mass of 380 grams, it was smaller than most modern pigeons. The most extraordinary feature of Yi qi is the styliform elongate bony rod projecting from the wrist: a long, pointed skeletal element that, together with a skin membrane (patagium), formed a wing of morphology entirely unlike any other known flying bird or dinosaur. This combination of a membrane wing supported by an accessory bony rod is unique among Dinosauria and finds functional parallels only in bats and flying squirrels among modern vertebrates. The holotype specimen (STM 31-2) was preserved with impressions of contour feathers on the body and filamentous plumage, plus portions of carbonized wing membrane visible around the forelimbs. Histological analysis of the feathers and bones indicates the animal had not reached full skeletal maturity at death, making adult size estimates slightly uncertain. Yi qi had long recurved foot claws consistent with arboreal habits, and simple conical teeth suggesting insectivorous or generalist carnivorous diet. The skull is relatively large in proportion to the body, with broad orbital openings, suggesting acute vision in a forest canopy environment. The phylogeny of Yi qi within Scansoriopterygidae is well-supported, but the placement of that clade within Coelurosauria remains debated: recent analyses alternate between positioning them as basal members of Pennaraptora or as an independent branch of theropods that developed gliding flight convergently and independently from modern birds. If Yi qi's patagium permitted active flight, it would represent a third independent origin of flight among feathered Jurassic dinosaurs, alongside the lineages leading to modern birds (Avialae) and possibly the gliding of Microraptor. Alternatively, aerodynamic analyses suggest the wing morphology was better suited to descending gliding from elevated perches than to sustained flapping. The discovery was announced in April 2015 in Nature by Xu Xing and collaborators, immediately becoming one of the most impactful paleontological finds of the decade.

Jurassic Carnivore 0.6m

Other Animals of the Mesozoic

Not dinosaurs, but they dominated the oceans, the skies, and parts of the continents during the same period.

Hybodus

Shark · Marine

Hybodus

"Curved tooth"

The most successful shark of the Mesozoic. Had two types of teeth: pointed for fish and flat for crustaceans — rare versatility. Shared the seas with ichthyosaurs, plesiosaurs, and mosasaurs.

Triassic–Cretaceous 2.5m Marine
Cretoxyrhina mantelli

Shark · Marine

Cretoxyrhina mantelli

"Sharp-snouted of the Cretaceous"

One of the largest sharks of the Cretaceous. Smooth unserrated teeth, high speed, morphology similar to the modern great white. Fossils show mosasaur vertebrae bearing its tooth marks.

Cretaceous 7m Marine
Squalicorax

Shark · Marine

Squalicorax

"Raven shark"

Shark with heavily serrated teeth similar to the modern tiger shark. Fossil evidence shows it fed on carcasses of dinosaurs that fell into rivers and seas — the open-ocean scavenger niche.

Cretaceous 5m Marine
Ichthyosaurs

Ichthyosaur · Marine

Ichthyosaurs

"Fish lizards"

Extraordinary convergence with dolphins and sharks, but they were reptiles that gave birth to live young at sea. Shonisaurus (Triassic) reached 21m. Ophthalmosaurus had the largest proportional eyes of any known vertebrate.

Triassic–Cretaceous 21m Marine
Plesiosaurs

Plesiosaur · Marine

Plesiosaurs

"Near lizard"

Elasmosaurus had ~14m of neck with 72 cervical vertebrae. They swam with four large flippers in a kind of "underwater flight," snatching individual fish with rapid neck strikes.

Jurassic–Cretaceous 14m Marine
Pliosaurs

Plesiosaur · Marine

Pliosaurs

"More lizard"

Sister group of plesiosaurs, but with a short neck and enormous head. Kronosaurus had a 2.7m skull. Predator X (Jurassic, Norway) may have reached 15m. Top predators of Jurassic oceans.

Jurassic–Cretaceous 15m Marine
Mosasaurus hoffmannii

Mosasaur · Marine

Mosasaurus hoffmannii

"Lizard of the Meuse River"

The largest known mosasaur at 17 meters. Related to monitor lizards and snakes, not crocodiles. First giant marine reptile fossil described by science (1764, Maastricht). Had a double-hinged jaw like snakes.

Cretaceous 17m Marine
Pterosaurs

Pterosaur · Aerial

Pterosaurs

"Winged lizards"

Not dinosaurs — they were the sister group. Had hair-like fibers (pycnofibers) and were likely endothermic. Quetzalcoatlus northropi: 10–11m wingspan, 2.5m tall on land. Largest flying animal of all time.

Triassic–Cretaceous 11m Aerial
Rauisuchids

Archosaur · Terrestrial

Rauisuchids

"Crocodilians of Rau"

While dinosaurs were still small, rauisuchids were the apex predators of the Triassic. Saurosuchus (7m), Postosuchus (6m), Fasolasuchus (8–10m). Their extinction cleared the way for large theropods.

Triassic 10m Terrestrial
Phytosaurs

Archosaur · Terrestrial

Phytosaurs

"Plant lizards"

Perfect convergence with modern crocodilians, but no close relation. Key difference: nostrils positioned almost between the eyes (in crocodilians they sit at the tip of the snout).

Triassic 6m Terrestrial
Crocodyliformes

Crocodylomorph · Terrestrial

Crocodyliformes

"Crocodile-shaped"

Far more diverse in the Mesozoic than today: bipedal terrestrial forms in the Triassic, pelagic marine forms in the Jurassic. Dakosaurus had a theropod-like skull. What we know today is just the survivors.

Triassic–Cretaceous 8m Terrestrial
Dicynodonts

Synapsid · Terrestrial

Dicynodonts

"Two dog teeth"

Synapsids with a horny beak and tusks. Lystrosaurus dominated the early Triassic after surviving the Permian mass extinction. Placerias (hippo-sized) briefly coexisted with the first dinosaurs.

Triassic 3m Terrestrial
Mesozoic mammals

Mammal · Terrestrial

Mesozoic mammals

"Mammals of the Age of Dinosaurs"

Small, likely nocturnal, living on the margins. But there was real diversity: Castorocauda (semi-aquatic, 164 Ma), Volaticotherium (glider, 160 Ma), Repenomamus (fossil found with a dinosaur hatchling in its stomach).

Triassic–Cretaceous 1m Terrestrial